Dictionary of Human Evolution and Biology

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1. evolution of two or more mutually advantageous adaptations in interactive species.

2. a process of harmonious integration of collaborating genes, such as the several genes involved in a sequential biochemical pathway; neutral mutations in coadapted systems may be suppressed by selection because so many dependent loci are affected, but an advantageous mutant at any one locus could permit optimizing mutations to cascade through the system as other loci accommodate to the original advantageous mutant.